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diff --git a/browser/themes/linux/browser.css b/browser/themes/linux/browser.css
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+/* This Source Code Form is subject to the terms of the Mozilla Public
+ * License, v. 2.0. If a copy of the MPL was not distributed with this
+ * file, You can obtain one at http://mozilla.org/MPL/2.0/. */
+
+@import url("chrome://browser/skin/browser-shared.css");
+@import url("chrome://browser/skin/contextmenu.css");
+
+@namespace html url("http://www.w3.org/1999/xhtml");
+
+/**
+ * We intentionally do not include browser-custom-colors.css, instead choosing to
+ * fall back to system colors and transparencies in order to accommodate the
+ * wider variety of GTK system themes.
+ *
+ * We still want to do some of the tweaks that browser-custom-colors does, like
+ * disabling the toolbar field border and backgrounds.
+ */
+@media not (prefers-contrast) {
+  :root:not(:-moz-lwtheme) {
+    --toolbar-field-border-color: transparent;
+
+    /* These colors don't exactly match the default dark color that buttons and
+     * textfields have in Adwaita[1][2], which would end up computing to
+     * rgba(0, 0, 0, .8 * .1) and rgba(255, 255, 255, 1 * .1) for light and
+     * dark, respectively.
+     *
+     * Instead, for the light theme we use .05 alpha, to increase the contrast
+     * of the text. For the dark theme, we use a darker background, which works
+     * because the toolbar background applies some white unconditionally, and
+     * matches what our default themes do in other platforms too.
+     *
+     * [1]: https://gitlab.gnome.org/GNOME/libadwaita/-/blob/42c04e038f19b2123560da662692d65480a67931/src/stylesheet/widgets/_buttons.scss#L1
+     * [2]: https://gitlab.gnome.org/GNOME/libadwaita/-/blob/42c04e038f19b2123560da662692d65480a67931/src/stylesheet/widgets/_entries.scss#L9
+     */
+    --toolbar-field-background-color: light-dark(rgba(0, 0, 0, .05), rgba(0, 0, 0, .3));
+    --toolbar-field-color: inherit;
+
+    @media (-moz-gtk-theme-family) {
+      --tabs-navbar-shadow-color: transparent;
+      @media (prefers-color-scheme: light) {
+        --urlbar-box-bgcolor: #fafafa;
+      }
+    }
+  }
+}
+
+#menubar-items {
+  flex-direction: column; /* for flex hack */
+  justify-content: normal; /* align the menubar to the top also in customize mode */
+}
+
+/**
+ * Titlebar drawing:
+ *
+ * GTK windows have both a window radius (exposed via the
+ * `-moz-gtk-csd-titlebar-radius`) environment variable, and a window shadow
+ * (which we can't read back from GTK). Note that the -moz-window-decorations
+ * (in X11) or the compositor (in Wayland) does draw the shadow corners
+ * already.
+ */
+:root[tabsintitlebar] {
+  background-color: transparent;
+
+  /* When temporarily showing the menu bar, make it at least as tall as the tab
+   * bar such that the window controls don't appear to move up. */
+  #toolbar-menubar[autohide="true"] {
+    height: calc(var(--tab-min-height) + 2 * var(--tab-block-margin));
+  }
+
+  &[sizemode="normal"]:not([gtktiledwindow="true"]) {
+    /* This takes care of drawing our window decorations on X11 */
+    -moz-default-appearance: -moz-window-decorations;
+    appearance: auto;
+
+    #navigator-toolbox,
+    dialog::backdrop {
+      border-top-left-radius: env(-moz-gtk-csd-titlebar-radius);
+      border-top-right-radius: env(-moz-gtk-csd-titlebar-radius);
+    }
+
+    /* stylelint-disable-next-line media-query-no-invalid */
+    @media (-moz-bool-pref: "widget.gtk.rounded-bottom-corners.enabled") {
+      #navigator-toolbox {
+        /* The body clip below covers this. */
+        border-radius: 0;
+      }
+
+      body,
+      dialog::backdrop {
+        /* Use an uniform clip to allow WebRender to optimize it better */
+        border-radius: env(-moz-gtk-csd-titlebar-radius);
+      }
+    }
+  }
+}
+
+/* Bookmark menus */
+.bookmark-item {
+  &:is(menu, menuitem) {
+    min-width: 0;
+    max-width: 32em;
+  }
+
+  &:not(.subviewbutton) > .menu-iconic-left {
+    margin-block: 0;
+  }
+
+  /* Bookmark drag and drop styles */
+  &[dragover-into="true"] {
+    background: SelectedItem !important;
+    color: SelectedItemText !important;
+  }
+
+  &[cutting] {
+    > .toolbarbutton-icon,
+    > .menu-iconic-left > .menu-iconic-icon {
+      opacity: 0.5;
+    }
+
+    > .toolbarbutton-text,
+    > .menu-iconic-left > .menu-iconic-text {
+      opacity: 0.7;
+    }
+  }
+}
+
+/* Address bar */
+#urlbar,
+#searchbar {
+  font-size: 1.05em;
+}
+
+/* stylelint-disable-next-line media-query-no-invalid */
+@media (-moz-bool-pref: "browser.urlbar.experimental.expandTextOnFocus") {
+  #urlbar[breakout-extend] {
+    font-size: 1.14em;
+  }
+}
+
+/* AutoComplete */
+
+#PopupAutoComplete > richlistbox > richlistitem[originaltype~="datalist-first"] {
+  border-top: 1px solid ThreeDShadow;
+}
+
+/* Content area */
+
+.sidebar-splitter {
+  appearance: none;
+  width: 6px;
+  background-color: -moz-dialog;
+  border: 1px ThreeDShadow;
+  border-style: none solid;
+}
+
+/* Tabstrip */
+
+#tabbrowser-tabs {
+  /* override the global style to allow the selected tab to be above the nav-bar */
+  z-index: auto;
+}
+
+#TabsToolbar {
+  min-height: 0;
+}
+
+#TabsToolbar,
+#toolbar-menubar,
+#main-menubar,
+#titlebar {
+  appearance: none;
+  background-color: transparent;
+  color: inherit;
+}
+
+/* All tabs menupopup */
+
+.alltabs-item[selected="true"] {
+  font-weight: bold;
+}
+
+#UITourHighlight {
+  /* Below are some fixes for people without an X compositor on Linux.
+     This is why we can't have nice things: */
+  /* Animations don't repaint properly without an X compositor. */
+  animation-name: none !important;
+  /* Opacity rounds to 0 or 1 on Linux without an X compositor, making the
+     background color not visible. Anti-aliasing is not available either. Make a
+     thicker outline and cancel border-radius for that case. */
+     outline: 4px solid rgb(0,200,215);
+     border-radius: 0 !important;
+}
+
+#UITourTooltipDescription {
+  font-size: 1.05rem;
+}
+
+#UITourTooltipClose {
+  margin-inline-end: -4px;
+}
+
+/**
+ * Override the --arrowpanel-padding so the background extends
+ * to the sides and bottom of the panel.
+ */
+#UITourTooltipButtons {
+  margin-inline-start: -10px;
+  margin-bottom: -10px;
+}
+
+/* The button box must appear on top of the navigator-toolbox in order for
+ * click and hover mouse events to work properly for the button in the restored
+ * window state. Otherwise, elements in the navigator-toolbox, like the menubar,
+ * can swallow those events. */
+.titlebar-buttonbox {
+  z-index: 1;
+  align-items: center;
+}
+
+/* When using lightweight themes, use our own buttons since native ones might
+ * assume a native background in order to be visible. */
+.titlebar-button:-moz-lwtheme {
+  appearance: none;
+  border-radius: 100%;
+
+  &:hover {
+    background-color: color-mix(in srgb, currentColor 12%, transparent);
+  }
+
+  &:hover:active {
+    background-color: color-mix(in srgb, currentColor 20%, transparent);
+  }
+
+  &:focus-visible {
+    outline: var(--focus-outline);
+    outline-offset: var(--focus-outline-inset);
+  }
+
+  > .toolbarbutton-icon {
+    padding: 6px;
+    -moz-context-properties: stroke;
+    stroke: currentColor;
+  }
+}
+
+/* Render titlebar command buttons according to system config.
+ * Use full scale icons here as the Gtk+ does. */
+.titlebar-max {
+  appearance: auto;
+  -moz-default-appearance: -moz-window-button-maximize;
+  order: env(-moz-gtk-csd-maximize-button-position);
+
+  &:-moz-lwtheme {
+    list-style-image: url(chrome://browser/skin/window-controls/maximize.svg);
+  }
+}
+
+.titlebar-restore {
+  appearance: auto;
+  -moz-default-appearance: -moz-window-button-restore;
+  order: env(-moz-gtk-csd-maximize-button-position);
+
+  &:-moz-lwtheme {
+    list-style-image: url(chrome://browser/skin/window-controls/restore.svg);
+  }
+}
+
+@media not (-moz-gtk-csd-minimize-button) {
+  .titlebar-min {
+    display: none;
+
+    &:-moz-lwtheme {
+      list-style-image: url(chrome://browser/skin/window-controls/restore.svg);
+    }
+  }
+}
+
+@media not (-moz-gtk-csd-maximize-button) {
+  .titlebar-restore,
+  .titlebar-max {
+    display: none;
+  }
+}
+
+.titlebar-close {
+  appearance: auto;
+  -moz-default-appearance: -moz-window-button-close;
+  order: env(-moz-gtk-csd-close-button-position);
+
+  &:-moz-lwtheme {
+    list-style-image: url(chrome://browser/skin/window-controls/close.svg);
+
+    &:hover {
+      background-color: #d70022;
+      color: white;
+    }
+
+    &:hover:active {
+      background-color: #ff0039;
+    }
+  }
+
+  @media not (-moz-gtk-csd-close-button) {
+    display: none;
+  }
+}
+
+.titlebar-min {
+  appearance: auto;
+  -moz-default-appearance: -moz-window-button-minimize;
+  order: env(-moz-gtk-csd-minimize-button-position);
+
+  &:-moz-lwtheme {
+    list-style-image: url(chrome://browser/skin/window-controls/minimize.svg);
+  }
+
+  @media not (-moz-gtk-csd-minimize-button) {
+    display: none;
+  }
+}
+
+@media (-moz-gtk-csd-reversed-placement) {
+  .titlebar-buttonbox-container {
+    order: -1;
+  }
+}